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. 2014 Apr 3;8(10):2104–2115. doi: 10.1038/ismej.2014.47

Table 1. Three organisational levels of BSCs as a functional unit, their traits, units and possible drivers or anticipated drivers of change.

Functional unit: BSC
Level Trait Unit Driver
BSC community members Lichen species Name Genetic limits to phenotypic/metabolic plasticity
  Number of species Number Overall metabolic fate: growth versus survival
  Functional group (mosses, lichens, cyanobacteria, green algae) Occurrence/number Succession state/habitat suitability
       
Carbon gain NP μmol m−2 s−1 Carbon gain
  Dark respiration μmol m−2 s−1 Maintenance/growth
  Carbon gain efficiency ratio Overall gain efficiency
  Light level to saturate photosynthesis μmol photons m−2 s−1 Light use/Protection against excess light
  Light level to achieve NP compensation μmol photons m−2 s−1 Low light use efficiency
  Thallus water content gH2O gDW−1 Water availability
  Optimal temperature for NP °C Active temperature
       
Carbon allocation Small sugars % Fixed carbon Stress protection/Respiratory substrate
  Polysaccharides % Fixed carbon Intermediate storage for growth support versus respiratory backup
  Insolubles % Fixed carbon Growth versus survival

Abbreviations: BSC, biological soil crust; NP, net photosynthesis.