The metabolism primarily occurs via cytochrome P450s (CYP1A1 and CYP1B1), peroxidases (epoxide hydrolase) and aldo-keto reductase. The major metabolite which is thought to cause carcinogenesis is BaP-7,8-diol-9,10-epoxide (BPDE) which is metabolised via BaP-7,8-epoxide and BaP-7,8-diol. BPDE forms DNA-adducts which can induce G→T transversions. In addition, many metabolites (such as catechol and dione within a futile cycle) form reactive oxygen species (ROS) thus adding to oxidative stress and contributing to the formation of oxidative DNA lesions such as 8-oxoguanine, a pre-mutagenic DNA lesion. Radical cations are formed in a minor quantity.