Table 4. Measured Distortions of the Zinc Binding Site^a.

	His15	His25	His85	Glu89
SC1982	95.5°	19.1°	0.7 Å	1.9 Å
SBi4172	76.9°		1.4 Å	2.5 Å
SBi4434		28.1°	67.9°	1.5 Å
SC1475	68.2°		0.8 Å	1.3 Å
SC124	82.9°		1.0 Å	1.6 Å
SC084435	38.3°	2.8 Å	157.7°	5.0 Å
SC032235	31.2°	31.7°	49.1°	not applicable

^aShown are averaged movements of side chains in angstroms (directional movement of C^γ) or degrees (swing of C^β–C^γ). As Glu89 was not modeled in 1MHO, directional measurements are from the carbonyl carbon atom of Phe88.