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. 2014 Oct 8;5:4979. doi: 10.1038/ncomms5979

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Anaphase I (a–c), telophase I (d), metaphase II (e,f) and anaphase II (g) chromosomes immunolabelled with centromere- (CENH3 or H2AThr120ph; red) and α-tubulin-specific (green) antibodies. The right images in a show enlarged spindle microtubules attachment along the entire length of the holocentric chromatids. Distribution of subterminal LeSAT7 (green) and telomere (red) repeats during anaphase I (b) and metaphase II (f), as well as of chromosome-specific markers (LeSAT7 (green) and LeSAT28 (red)) in tetrads (h) after FISH. The insets and arrows in b,f show LeSAT7-enriched chromatin threads connecting two homologous non-sister chromatids. Equal number of LeSAT7 and LeSAT28 signals in each tetrad nucleus indicates proper haploidization (six signals of LeSAT7 and two signals of LeSAT28 per tetrad nucleus). DNA counterstained with DAPI (blue). Scale bars, 10 μm.