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Boundary elements STAR and TEF2-UAS block URA3 silencing in the presence of a clone-601 mononucleosome. Clone-601-167×1 DNA (167 bp NRL, open arrow) was positioned between the HML–E silencer and the URA3 reporter gene. The antisilencers STAR (A and B) or TEF2-UAS (C and D), shown by shaded arrows, were positioned upstream of the HML–E silencer in different orientations. The URA3 reporter status was tested by spotting assay as in Figure 1.
