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Identification of drugs altering PHT1;4 expression. A, Screen used to identify PTN molecules. GUS staining of pht1;4-1 plantlets germinated and grown 5 d on +Pi (500 µm) and transferred for 5 d on +Pi, –Pi (15 µm), or –Pi plus PTN1 (25 µm). B, Chemical structure of PTN1.