Table 2. Estimates of components of variation results of the four-way PERMANOVA on single factors and interaction terms for all analyzed variables.

Single factor and/or interaction terms	MGT	MEGD	MEMD	MJD	MALS	OS
S	2.893	0.526	−0.004	0.870	0.637	183.480
Sp	7.217	0.002	0.311	0.748	3.776	452.880
T	169.230	25.561	10.500	14.894	124.740	93.748
Sal	9.277	1.634	0.106	0.455	0.230	71.800
T x Sal	0.223	0.005	0.046	0.371	0.080	30.517
Sp x Sal	3.163	1.484	0.039	0.197	0.201	25.923
Sp x T	0.384	0.010	0.050	0.101	0.144	3.972
S x Sp	5.498	0.039	2.793	1.826	12.950	160.550
S x Sal	1.744	0.020	0.016	0.214	0.090	13.748
S x T	1.931	0.768	0.005	0.061	0.482	12.609
S x T x Sal	0.139	0.126	0.213	−0.031	0.474	142.200
S x Sp x T	0.210	−0.007	0.026	0.072	0.176	7.579
Sp x T x Sal	0.790	0.090	0.021	0.189	0.082	13.487
S x Sp x Sal	0.232	0.070	0.073	0.151	0.077	20.431
S x Sp x T x Sal	0.696	0.130	0.550	0.197	0.024	44.797
V(Res)	0.269	0.157	0.356	0.432	0.451	20.371

The components of variation was estimated by PERMANOVA (Euclidean distances, 9999 permutations) for MGT, minimum generation time; MEGD, minimum time until first egg(s) deposition; MEMD, minimum time for embryonic development; MJD, minimum time for the development of juveniles into adults; MALS, minimum adult life span; OS, offspring per female at the end of the experiment. High values correspond to a high relative importance of the respective factor.

Abbreviations: S, absence/presence of sulphide; T, temperature (16, 10 and 4°C); Sp, cryptic Halomonhystera disjuncta species GD1–3 (previously named Geomonhystera disjuncta); Sal, salinity (25, 29.5 and 34 psu).