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A hydrogen OMIT map illustrating the protonation state of His57: N∊ is clearly protonated pointing towards Ile59, leaving the Nδ atom unprotonated and allowing it to act as a hydrogen-bond acceptor of terminal hydroxyl groups. The electron-density map (2F o − F c) contoured at the 2.5σ level is coloured blue and the difference density map (F o − F c) is coloured green. Protein residues and the uridine nucleotide are shown in ball-and-stick representation.
