Schematic representation of development of seedlings of H. courbaril growing in greenhouse, growth room and forest environments. Data represent the mean emergence dates of (A) seedlings, (B) eophylls, and (C) first, (D) second and (E) third metaphylls of seedlings with and without cotyledons. The detachment of cotyledons was performed at 22, 24 and 27 days after the beginning of imbibition (black arrows). Asterisks mark the date by which 50 % of cotyledons had dropped.