Table 3. Quantitative trait loci (QTLs) for threshability and brittle rachis identified in the Triticum aestivum ssp. tibetanum Shao population from accessions Q1028×ZM9023 by using the inclusive composite interval mapping (ICIM) analysis.

Trial type	QTL	Chromosomea	Trial	Positionb	Left Marker	Right Marker	LOD	PVE (%)	Additive effect
Brittle Rachis	Qbr.sau-2D1	2DS	RF2012	56	Xgpw361	Xgdm35	3.3	6.6	1.4
			RF2013	56	Xgpw361	Xgdm35	2.5	5.1	1.3
			RF combined	56	Xgpw361	Xgdm35	2.6	5.3	1.3
	Qbr.sau-2D2	2DL	RF2012	197	wPt-666518	wPt-730613	8.1	18.2	2.4
			RF2013	197	wPt-666518	wPt-730613	8.1	18.4	2.5
			RF combined	197	wPt-666518	wPt-730613	8.2	18.6	2.4
	Qbr.sau-3D	3DS	RF2012	112	wPt-9258	Xgdm72	16.6	39.3	3.5
			RF2013	112	wPt-9258	Xgdm72	15.0	36.9	3.5
			RF combined	112	wPt-9258	Xgdm72	16.1	38.7	3.5
	Qbr.sau-5A	5AL	RF2012	117	Xgpw2059	Xgpw4457	7.4	16.7	2.2
			RF2013	116	Xgpw2059	Xgpw4457	8.1	18.9	2.5
			RF combined	116	Xgpw2059	Xgpw4457	7.9	18.6	2.4
Threshability	Qft.sau-2D1	2DS	FT2012	68	Xcfd53	Xgpw4080	8.3	17.5	−9.5
			FT2013	67	Xcfd53	Xgpw4080	7.9	16.2	−10.7
			FT combined	68	Xcfd53	Xgpw4080	8.4	17.4	−10.1
	Qft.sau-2D2	2DL	FT2012	195	wPt-666518	wPt-730613	6.1	13.1	−8.3
			FT2013	195	wPt-666518	wPt-730613	5.9	12.5	−9.5
			FT combined	195	wPt-666518	wPt-730613	6.3	13.2	−8.9
	Qft.sau-5A	5AL	FT2012	121	Xgpw4457	Xgpw2273	14.1	38.3	−14.2
			FT2013	122	Xgpw4457	Xgpw2273	11.1	29.9	−14.5
			FT combined	122	Xgpw4457	Xgpw2273	12.9	35.2	−14.4

LOD: limit of detection; PVE: phenotypic variation explained.

^aMarkers on the chromosome arm are considered to be linked with the QTL.

^bThe position of LOD score peak in the chromosomes (Figs. 4–6).