Table 1. Description of state variables.

State variable	Description
Sh	Number of humans who are not infected with malaria but may get infected if bitten by infectious mosquitoes.
Eh	Number of humans that are infected with malaria, but not yet infectious
Ih	Number of humans that are fully infectious for malaria
Rh	Number of humans in the recovered state who are immune to clinical disease but still harbour low levels of parasites and are still infectious to mosquitoes (albeit at a lower probability)
W	Mean W. bancrofti adult worm burden in the human population
F	Mean microfilariae burden in the human population
Mss	Number of mosquitoes uninfected with either parasite
Mse	Number of mosquitoes exposed to W. bancrofti, but not infected with P. falciparum
Msi	Number of mosquitoes with infectious stage W. bancrofti larvae, but not infected with P. falciparum
Mes	Number of mosquitoes exposed to P. falciparum, but not infected with W. bancrofti
Mee	Number of mosquitoes exposed to P. falciparum and W. bancrofti
Mei	Number of mosquitoes exposed to P. falciparum, and infectious for W. bancrofti
Mis	Number of mosquitoes with P. falciparum sporozoites, but not infected with W. bancrofti
Mie	Number of mosquitoes with P. falciparum sporozoites, and exposed to W. bancrofti
Mii	Number of mosquitoes infectious for both P. falciparum and W. bancrofti