Table 1. Average Spearman rank correlation coefficients <R> across all 50 LODs between all applied measures and fitness.

		<#concepts>	<ΣφMax>	<#MC elements>	<#MC concepts>	<ΦMax>
Task 1	<R>	0.38	0.28	0.13	0.14	0.11
	SEM	0.05	0.05	0.05	0.05	0.05
Task 2	<R>	0.55	0.48	0.22	0.22	0.21
	SEM	0.04	0.04	0.05	0.05	0.05
Task 3	<R>	0.47	0.39	0.30	0.28	0.27
	SEM	0.04	0.04	0.05	0.05	0.05
Task 4	<R>	0.71	0.63	0.47	0.48	0.50
	SEM	0.04	0.03	0.06	0.06	0.06

Since the average number of concepts and their Σφ^Max values capture both modular and integrated causal relations in the animat's brain as a whole, they correlated more strongly with fitness than the average number of MC elements, MC concepts and <Φ ^Max>, which nevertheless increased with adaptation. See S1 Fig. for complementary histograms of the correlation coefficients of all individual LODs.