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Pathways for the oxidation of benzene (0.8 mM) to phenol, for the oxidation of phenol (0.8 mM) to dihydroxybenzenes, and for the oxidation of dihydroxybenzenes (0.8 mM) to trihydroxybenzenes by E. coli TG1/pBS(Kan)ToMO expressing wild-type ToMO and the I100Q, F205G, and M180T/E284G TouA variants. The molar product percentages are indicated; the values in boldface type in parentheses indicate the product formation rates (in nanomoles per minute per milligram of protein).
