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H4 forms triple-helices. (A) CD spectrum shows positive MRE (mean residue ellipticity) band at 225 nm, consistent with triple-helical structure. (B) Thermal denaturation profile shows cooperative unfolding of H4 with a Tm of 33 °C. (C) WAXS of H4 shows interhelical d-spacing consistent with fiber packing observed in natural collagens.
