Figure 8. Examples of movement tracks for a starved Pterostichus melanarius beetle with low motility (7 cm² s⁻¹; left) and for a fed beetle with a high motility (88 cm² s⁻¹; right).

The total time spent in the arena's interior was 24.8 min for the starved and 11.7 min for the fed beetle. Moves in the 10 cm edge zone are represented by thin grey lines.