Table 1.

Mean ± SD relatedness estimates. Relatedness between all individuals, males and females (a) at the level of population and colony, and (b) at the level of social unit within colony [within Girvan–Newman partitions (‘within’) or between Girvan–Newman partitions (‘between’) in the same colony].

(a)	Population	Colony
All (1138)	−0.001 ± 0.151	0.032 ± 0.175***
Male (561)	0.000 ± 0.150	0.055 ± 0.191***
Female (530)	−0.002 ± 0.149	0.019 ± 0.162**

	Girvan–Newman partitions
(b)	Within	Between
All (314)	0.054 ± 0.190	0.025 ± 0.169***
Male (174)	0.120 ± 0.213	0.036 ± 0.182***
Female (119)	0.035 ± 0.161	0.015 ± 0.156n.s.

Comparisons between mean relatedness of the entire population and those at the level of colony were made using one-sample Wilcoxon singed rank tests with the mean relatedness at the population level set as μ. Numbers in parentheses indicate samples size of the genotyped population. See text for statistics of Girvan–Newman partitions.

^**P < 0.01

^***P < 0.001; n.s., not significant.