Wab1 acts in a lateral domain in the maize leaf. A, Leaf founder cells are initially recruited into central (C, blue), lateral (L, red), and marginal (M, green) domains in the SAM; the marginal domain is not recruited in ns mutants. B, During P1 to P2, growth of the lateral domain is inhibited in Wab1 mutants, resulting in a narrow primordium. C, Cells in the lateral domain contribute the majority of the distal blade tissue in the mature leaf and this tissue is reduced in Wab1 leaves. Cells in the margin domain contribute the majority of the sheath and proximal blade tissue in the mature leaf and this tissue is absent in ns leaves. Wab1;ns mutants reveal both defects.