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. 2014 Dec 17;27(2):172–197. doi: 10.1007/s12640-014-9508-6

Table 4.

Readers of epigenetic modifications

Recognition site Reader domain Protein Modification
Methylcytosine MBD MBD1, MBD2, MBD4, MeCP2 5-methylcytosine (5mC)
SRS UHRF1, UHRT2
BTP/POZ-Zn-finger Kasio, ZBTB4, ZBTB38
IPT/TIG RBP-J
Methyllysine Chromobarrel (R) MOF, Eaf3, MRG15, H3K36me2/me3, H3K4me1, H4K20me1
CG (R) HP1, CDH1, PC, MPP8, CDY, CDYL, CDYL2, CBX7, MCL3, H3K9me2/me3, H3K27me2/me3
Tudor (R) PHF1, PHF19, PHF20, TDRD7, H3K36me3
TTD (R) 53BP1, KDM4A, KDM4B, KDM4C, Sgf29, UHRF1 H3K4me3, H3K9me3, H4K20me2
MBT (R) CGI-73, L(3)HBTL, SFMBT, PHF20L1 H3K4me1, H3K9me1/me2, H3K20me1, H4K4me1
PWWP (R) DNMT3A, BRPF1, NSD1-3, MSH-6, N-PAC, Pdp1 H3K36me3, H4K20me1/me3, H3K79me3
ADD (Pt) DNMT3L H3K9me3
Ankyrin repeats G9a/GLP H3K9 me1/me2
BAH ORC1 H4K20me2
DCD (R) CHD1 H3K4me1/me2/me3
PHD (Pt) BHC80, BPTF, AIRE, RAG2, ING1-5, BPTF, TAF3, PHF2, PNF8,PHF13, PHF13, Pygo, YNG1, SMCX, H3K4me3, H3K4me2, H3K9me3
WD40 WDR5/WDR9, EED, LRWD1, H3K27me3, H3K9me3
zf-CW (Pt) ZCWPW1 H3K4me3
Methylarginine ADD (Pt) DNMT3L H4R3me2 s
Tudor (R) AKAPI1, TDRD2-3, TDRD5,TDRD8-10, SMN1, SPF30, H3Rme2, H4Rme2
WD40 WDR5 H3R2me2
Acetyllysine BD GCN5, PRBM1, H3Kac, H4Kac, H2AKac, H2BKac
DBD (R) Rsc4, TAF1, Brdt, H3KacKac, H4KacKac
DFP (Pt) DPF3b H3K14ac
PH histone chaperone Rtt106 H3K56ac
Phosphoserine 14-3-3 14-3-3ξ, 14-3-3β, 14-3-3γ, 14-3-3η, 14-3-3ε, 14-3-3μ, 14-3-3θ H3S10ph, H3S28ph
tandem BRCT H2A.XS139 (γH2AX)
Phosphothreonine BIR H3T3ph
Propionyllysine BD Brd4 H3K23
Butyryllysine BD Brd4 K3K14
Unmodified H3 ADD (Pt) DNMT3L unmodified histone H3
PHD (Pt) UHRF1 unmodified histone H3
WD40 Nurf55 unmodified histone H3

ADD ATRX-DNMT3-DNMT3L, BAH bromo-adjacent homology, BD bromodomain, CD chromodomain, DCD double chromodomain, DBD double bromodomain, DFP double PHD finger, MBD methyl-CpG-binding domain, MBT malignant brain tumor, PH double pleckstrin homology, PHD plant homeodomain, PWWP Pro-Trp-Trp-Pro; SRA SET- and Ring finger-associated domain, TTD tandem Tudor domain, zf-CW zinc finger CW, (Pt) PHD-type, (R) Royal superfamily