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. 2015 Feb 6;9:10. doi: 10.3389/fncom.2015.00010

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Copyright © 2015 Page, Walters and Stringer.

This is an open-access article distributed under the terms of the Creative Commons Attribution License (CC BY). The use, distribution or reproduction in other forums is permitted, provided the original author(s) or licensor are credited and that the original publication in this journal is cited, in accordance with accepted academic practice. No use, distribution or reproduction is permitted which does not comply with these terms.

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Hypothesized path integration mechanism (left) and resulting firing times of HD cells (right). Cell 1 projects most strongly to cell 2, with delay Δt. If cell 1 is active at time t, cell 2 will therefore become active at time t + Δt. The angular separation between cells 1 and 2 will therefore govern the observed activity packet speed i.e., how far the activity packet has moved after a delay of Δt.