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A phylogeny of the Nymphalidae showing all tribes (and subtribes within the tribe Satyrini), modified from [ 17 ]. The clades analyzed in this study are marked by an eyespot; because Parargina, Mycalesina, and Lethina form a clade, they were analyzed together, as indicated by the eyespot at the node shared by these three subtribes. Colored branches represent different subfamilies.
