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. Author manuscript; available in PMC: 2015 Oct 1.
Published in final edited form as: J Neuroendocrinol. 2014 Oct;26(10):665–684. doi: 10.1111/jne.12183

Table 1.

Functional magnetic resonance imaging (fMRI) Studies of Mothers Highlighting Contrasting Findings in Healthy Mothers Versus Those With Affective Symptoms, Substance Use, or Characteristics Relating to Impaired Attachment Processes.

Author (year) (reference) N; Group Age of infants/
children
fMRI paradigm Findings
Salience/fear network
  Healthy mothers
    Seifritz et al. (2003) (40) 20, mothers versus
women without
children
< 3 years Infant cries and laughter
versus control sound
↑ Amygdala, middle cingulate
cortex and insula activity in
mothers versus women without
children
    Bartels and Zeki (2004) (45) 19, mothers 9 months to
6 years
Face pictures of own versus
unknown/familiar child;
comparison of maternal
versus romantic love
↑ Insula and dACC activity to own
child, which overlapped regions
active to romantic others
    Ranote et al. (2004) (101) 10, mothers 4–8 months Video of own versus other
infant
↑ Amygdala activation to own
infant
    Swain et al. (2008) (42) 12, vaginal versus
caesarean section
deliveries
2–4 weeks infantcry (own versus other) ↑ Amygdala activity to infant cry
in women with vaginal versus
caesarean section deliveries
    Lenzi et al. (2009) (114) 16, primiparous
mothers
6–12 months Viewing and imitating faces of
own versus other child
↑ Insula, amygdala and inferior
frontal gyrus activity associated
with increased maternal
empathy
    Strathearn et al. (2013) 39, primiparous
mothers
~11 months Child faces (own versus other) ↑ Amygdala activity to own
versus other infant faces
    Ho et al. (2014) (216) 14, mothers ~4 years Parenting decision-making
task
↑ vACC to +ve versus −ve infant
feedback. Hypothalamus-septa
coupling during −ve feedback
inversely related to cortiso
reactivity
  Mothers with affective symptoms, substance use or impaired infant interaction
    Silverman et al. (2007) (49) 8, postpartum mothers
with EPDS > 12 or
<6
7–8 weeks Negative emotional words ↓ Amygdala but ↑ insula activity
to negative words in women
with EPDS > 12
    Moses-Kolko et al. (2010) (50) 30, PPD versus HC 8–10 weeks Fear/threat adult faces ↓ DMPFC activity and amygdala-
DMPFC connectivity in PPD
↓ Amygdala activity associated
with postpartum depression
severity
    Laurent & Ablow (2012) (43) 22, PDD versus HC 18 months infantcry (own versus other) ↓ dACC and insula activity in PPD
    Barrett et al. (2012) (39) 22, mothers, varied
affective symptoms
~3 months Infant faces (own versus
other), with affect rating
task
↓ Amygdala activation to positive
own infant faces associated with
more depression (EPDS) and
anxiety symptoms (STAI-trait)
    Schechter et al. (2012) (61) 20, IPV-PTSD versus
control
12–42 months Videos of child stress during
separation (own versus other
child)
↑ Insula and ↓ mPFC activity to
child separation videos that
correlated with self-reported
stress
    Moser et al. (2013) (63) 20, IPV-PTSD 12–42 months Videos of child stress with
separation (own versus other
child)
↓ Cingulate, hippocampus and ↑
DLPFC activity to child
separation videos associated
with greater dissociative
symptoms
    Landi et al. (2011) (128) 54, substance using
versus non-using
postpartum mothers
1–3 months Infant sad faces, low distress
cries (all other infants)
↓ Amygdala, parahippocampus,
insula, lateral PFC, activity in
substance users
    Musser et al. (2012) (157) 22, primiparous, varied
maternal sensitivity
18 months infantcry (own versus other
versus control sounds)
↓ IFG and ↑ insula activity
associated with reduced
maternal sensitivity and
increased maternal intrusiveness
    Laurent et al. (2012) (156) 22, primiparous
mothers
15–18 months infantcry (own versus control
sound)
↑ Parahippocampus/amygdala and
insula activity to cry in mothers
of insecurely-attached infants

Author (year) (reference) N; Group Age of infants/
children
Paradigm, variable Regional findings

Executive network
  Healthy postpartum versus menstruating nonpostpartum women
    Bannbers et al. (2013) (76) 26; postpartum versus
menstruating women
48 h; 4–7 weeks Go/NoGo task ↓ VLPFC, precentral gyrus, and
dACC at later versus early
postpartum timepoints and
relative to nonpostpartum
women
Reward/social attachment network
  Healthy mothers
    Lorberbaum et al. (2002) (41) 10, mothers 1–2 months infantcry (other) versus white
noise versus baseline rest
↑ Medial thalamus, medial PFC,
right OFC to infant cry
    Bartels and Zeki (2004) (45) 19, mothers 9 months–6 years Face pictures of own versus
unknown/familiar child
↑ Striatum, thalamus, PAG to own
child
    Leibenluft et al. (2004) (44) 7, mothers 5–12 years Child faces (own versus
familiar/unfamiliar other)
↑ Mentalising circuitry (DMPFC,
MPFC, PCC, TPJ, pSTS) to own
child > familiar/unknown child
    Noriuchi et al. (2008) (37) 13, mothers 15–20 months Video clips own versus other
infant
↑ OFC, periaqueductal gray,
putamen to own infant
    Strathearn et at. (2008) (38) 26, primiparous
mothers
7–10 months Child happy faces (own versus
other)
↑ Putamen, SN, thalamus,
amygdala to own versus other
faces
    Swain et al. (2008) (42) 12, vaginal versus
caesarean section
deliveries (VD)
2–4 weeks infantcry (own versus other) ↑ SFG, caudate and thalamus to
infant cry in women with
vaginal versus caesarean section
deliveries. Correlation of parental
preoccupation with lenticular
nucleus activation
    Kim et al. (2011) (48) 17, breast- versus
formula-feeding
mothers
2–4 weeks and
3–4 months
infantcry (own versus other) ↑ SFG, insula, precuneus, striatum,
and amygdala activity to own
infant cry in breast- versus
formula-feeding mothers
↑ Maternal sensitivity associated
with increased SFG and
amygdala activity
  Mothers with affective symptom, impaired infant interaction, or low maternal care received, insecure attachment
    Silverman et al. (2007) (49) 8, postpartum mothers
with EPDS > 12 or
<6
7–8 weeks Positive emotional words ↓ Striatum to positive words in
women with EPDS > 12
    Strathearn et al. (2009) (46) 30, primiparous
mothers
~11 months Child happy faces (own versus
other)
↓ VST, OFC, mPFC activity to faces
in mothers without secure
attachment
    Kim et al. (2010) (53) 26, mothers 2–4 weeks infantcry (other versus
control sound)
↓ MFG, STS and fusiform gyrus to
infant cry for women with a
history of lower maternal care
received
    Moses-Kolko et al.
(2011) (105)
24, PPD versus HC 8–10 weeks Monetary reward task ↓ Ventral striatal duration of
reward-related response in PPD
    Laurent & Ablow (2011) (43) 22, PDD versus HC 18 months infantcry (own versus control
sound)
↓ Thalamus, caudate, OFC, SFG in
PPD, and in association with
higher CESD scores
    Atzil et al. (2011) (125) 23, mothers 4–6 months Mother–infant interaction
videos (self versus other
mother–infant dyad)
↑ Striatal activity associated with
greater mother–infant behavioral
synchrony
Default mode network
  Depressed versus healthy mothers
    Xiao-juan et al. (2011) (147) 21, PPD versus HC Within 16 weeks Resting state ↑ PCC index of nearest neighbor
connectivity (ReHo) in PPD
    Chase et al. (2013) (144) 37, PPD versus HC 10 weeks Resting state ↓ PCC-amygdala connectivity in
PPD
    Deligiannidis et al.
(2013) (146)
17, PPD versus HC 9 weeks Resting state ↓ Connectivity within
corticolimbic circuits including
the default mode network

CESD, Center for Epidemiologic Studies Depression Scale; dACC, dorsal anterior cingulate cortex; DMPFC, dorsomedial prefrontal cortex; EPDS, Edinburgh Postnatal Scale for Depression; HC, healthy controls; IFG, inferior frontal gyrus; IPV-PTSD, interpersonal violence-related post-traumatic stress disorder; MPFC, medial prefrontal cortex; OFC, orbitofrontal cortex; OT, oxytocin; PAG, periaqueductal gray; PCC, posterior cingulate cortex; PPD, postpartum depression; pSTS, posterior superior temporal sulcus; ReHo, regional homogeneity or synchronisation between time series of a voxel with that of its nearest neighbors; SFG, superior frontal gyrus; SN, substantia nigra; STAI, Speilberger State Trait Anxiety Inventory; TPJ, temporal–parietal junction; VE, valence. References are listed chronologically. In cases where references have findings that relate to multiple networks, such references have been categorised according to the most relevant network and, in some cases, are listed in two categories if more than one network is adequately assessed.