Female wing coloration and male sexual responses. Photographs females of four Hetaerina species: (a) H. cruentata (mating), (b) H. americana (marked for identification), (c) H. occisa, (d) H. titia. Sample reflectance spectra of female wings (e), with line colours matching the frames of the respective species' photographs (a–d). Wing lightness (f) affects whether H. titia females elicit a sexual response (stars) or not (circles) from H. americana (two-sided Mann–Whitney test, n = 14, p = 0.01) and H. occisa males (n = 77, p < 0.0001). (Online version in colour.)