Mean annual cycles of key ecosystem fluxes for three time periods of the fully forced C−N case. (A) GPP, (B) net N mineralization, (C) net ecosystem exchange (NEE, positive = CO2 source), and (D) heterotrophic respiration. Relative increase in GPP between experiments is smaller than proportional increase in N mineralization with deeper decomposition. Shift in N mineralization with enhanced deeper SOM decomposition toward autumn is due to longer decomposing than growing seasons, and phase lag of temperature in deep soils. The solid and dashed lines represent Zτ = 0.5 m and 10 m, respectively. All cases show the mean of the geographic region in which permafrost initially occurs in the model.