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Nucleosome asymmetry in vivo. (a) Assembly of asymmetric H3/H4 tetramers using a tandem affinity tag strategy. (b) Distribution of H3K27 methyl marks in ES cells into symmetric and asymmetric mononucleosomes. (c) Bivalent domains consist of asymmetric nucleosomes with one H3 tail di- or trimethylated at Lys4, and another tail marked with K27me2/3.
