Table 2.
The 43 morphological characteristics extracted by L-measure software and their descriptions.
| Number | Abbr. | Morphological index | Description |
|---|---|---|---|
| 1 | SS | Soma_surface | Somatic surface area |
| 2 | N s | N_stems | Total number of trees |
| 3 | N bi | N_bifs | Total number of bifurcations |
| 4 | N br | N_branch | Number of bifurcations plus terminations |
| 5 | N t | N_tips | Number of terminal tips of a neuron |
| 6 | NW | Neuronal_width | 95% of second principal component |
| 7 | NH | Neuronal_height | 95% of first principal component |
| 8 | ND | Neuronal_depth | 95% of third principal component |
| 9 | Ty | Type | Compartments are assigned to four different types: 1 = soma, 2 = axon, 3 = dendrites, and 4 = apical dendrites |
| 10 | Di | Diameter | Average branch diameter |
| 11 | Dp | Diameter_pow | Diameter of each compartment of the neuron raised to the power of 1.5 |
| 12 | Le | Length | Total arborization length |
| 13 | Su | Surface | Surface area of each compartment |
| 14 | SA | Section area | Total arborization surface area |
| 15 | Vo | Volume | Total internal volume of the arborization |
| 16 | ED | Euc distance | Maximum euclidean (straight) distance from soma to tips |
| 17 | PD | Path distance | Maximum path (along the tree) distance from soma to tips |
| 18 | BO | Branch_order | Maximum branch order number of bifurcations from soma to tips |
| 19 | Td | Terminal degree | Total number of tips each segment will terminate into |
| 20 | TS | Terminal segment | Number of compartments that comprise the terminal branch |
| 21 | Ta1 | Taper_1 | The change in diameter over path length between two critical points |
| 22 | Ta2 | Taper_2 | The ratio of the change in diameter to the initial diameter of two critical points. The initial diameter is usually larger |
| 23 | Bpl | Branch_path length | Summation of the individual compartment lengths that form a branch |
| 24 | Co | Contraction | Average contraction (the ratio between euclidean and path length calculated on each branch) |
| 25 | Fr | Fragmentation | Total number of reconstruction points |
| 26 | DR | Daughter_ratio | Ratio between the diameter of the bigger daughter and the smaller daughter of the current bifurcation |
| 27 | PDR | Parent-daughter_ratio | Ratio between the diameter of a daughter and its father for each critical point |
| 28 | Pa | Partition_asymmetry | Average over all bifurcations of the absolute value of (n1 − n2)/(n1 + n2 − 2), where n1 and n2 are the numbers of tips in the two subtrees |
| 29 | RP | Rall_power | Average over all bifurcations of the sum of the diameters of the two daughters, elevated to 1.5, divided by the diameter of the parent, and elevated to 1.5 |
| 30 | Pk | Pk | n = [0, 5] |
| 31 | Pc | Pk_classic | Rall power is set to 1.5 |
| 32 | Pk2 | Pk_2 | Rall power is set to 2 |
| 33 | Bal | Bif_ampl_local | Average over all bifurcations of the angle between the first two daughter compartments |
| 34 | Bar | Bif_ampl_remote | Average over all bifurcations of the angle between the following bifurcations or tips |
| 35 | Btl | Bif_tilt_local | The angles between the end of the parent branch and the initial part of the daughter branches at the bifurcation |
| 36 | Btr | Bif_tilt_remote | The angles between the previous node of the current bifurcating father and the daughter nodes |
| 37 | Btol | Bif_torque_local | Angle between the current plane of bifurcation and the previous plane of bifurcation |
| 38 | Btor | Bif_torque_remote | Angle between the current plane of bifurcation and the previous plane of bifurcation |
| 39 | Lpd | Last_parent_diam | Diameter of last bifurcation before the terminal tips |
| 40 | Dt | Diam_threshold | Diameter of first compartment after the terminal bifurcation leading to a terminal tip |
| 41 | HT | Hillman threshold | Computation of the weighted average diameter between 50% of father and 25% of daughter diameters of the terminal bifurcation |
| 42 | He | Helix | Helicity of the branches of the neuronal tree. It needs to be at least 3 compartments long to compute the helicity |
| 43 | FD | Fractal_dim | Fractal dimension metric of the branches in the dendrite trees |