Quantitative Trait Loci (QTL) identified for several traits under variable conditions on the three of the seven Linkage Groups (LG) of this study. SD; short day, LD ; long day, OE : outdoor environment, CA : cold acclimation, FT : freezing treatment, 1DF : 1 day post freezing, MF : mild freezing (Jan., 2012), SF : severe freezing (Feb., 2012).