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Biosynthetic pathway for tRNA s2U formation in E. coli (black arrows) and proposed pathway in B. subtilis (blue arrows). (Inset) Position of the tRNAGlu, tRNAGln, and tRNALys wobble uridine (U34) and structure of modified 2-thiouridine. The R group at position 5 of U34 can occur as H, CH2NH2, CH2NHCH3, CH2NHCH2CO2−, or CH2CO2CH3.
