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Genetic and phenotypic variation in Rev over time. The order of appearance of high frequency Rev mutations was determined by the day each mutant first dominated (frequency >50 %) the sampled population. Rev nuclear-export activity is reported relative to R1. Variants that differed significantly from R1 are indicated by asterisks: *P<0.05, **P<0.005, ***P<0.0005. #, Indicates significant difference from the prior ancestral sequence.
