Stride-related activity of 12 neurons with cutaneous RFs restricted to the ventral side of the forepaw. A, Vertical ground reaction forces during walking on the flat surface (solid line) and horizontal ladder (interrupted line). SDs were similar between the tasks and for clarity are shown only for flat surface walking (error bars). Stars indicate statistically significant differences between the walking tasks (p < 0.05, t test). B, Activity profiles of the 12 cells during locomotion on the flat surface. Vertical lines indicate the moment of the foot making contact with the ground (red) and 15 ms later (green). C, Average activity profile of the 12 cells, each normalized by its mean firing rate during flat surface walking.