Table 3. Summary of putative Early Cretaceous non-titanosauriform macronarians and other broad-toothed sauropods, according to previous studies.
Taxon | Formation/Age/Country | Identification |
---|---|---|
* Aragosaurus | Villar del Arzobispo Formation (late Tithonian-middle Berriasian), Spain | Identified as a titanosauriform by Canudo et al. [152] and D’Emic [17], but these opinions were not based on inclusion of Aragosaurus into a phylogenetic analysis. Recovered as MN in all analyses to date (e.g. [18,80,109,111]; LSDM and LCDM here). |
Cedarosaurus | Cedar Mountain Formation (Barremian-early Albian), USA | Recovered as MN by Royo-Torres [109, but most other analyses place this taxon as a brachiosaurid [1,17–19], LCDM here) or somphospondylan ([122], CSM here). |
Chubutisaurus | Cerro Barcino Formation (Aptian-Cenomanian), Argentina | Recovered as a MN by Carballido et al. [15], but most other analyses place this taxon as a somphospondylan [14,17–19], CSM here) or at least a titanosauriform (LCDM here). |
Dongbeititan | Yixian Formation (Barremian), China | Recovered as MN in the LCDM analysis of Mannion et al. [18], but this is not supported by their LSDM analysis or the LCDM result here. |
Euhelopus | Mengyin Formation (Barremian-Aptian), China | Recovered as MN by Carballido et al. [15], Carballido and Sander [19] and CSM Here, but most other analyses place this taxon within Somphospondyli ([10,13,17,18], LSDM and LCDM here) or at least a titanosauriform [14] |
* Galveosaurus | Villar del Arzobispo Formation (late Tithonian-middle Berriasian), Spain | Recovered as EU by Royo-Torres and Upchurch [110] in part, and Royo-Torres et al. [111], but most studies support an MN position ([15,18,19], CSM, LSDM and LCDM here). |
* Haestasaurus | Hastings Beds Group (late Berriasian-Valanginian), UK | Regarded as a titanosauriform [17], somphospondylan [18,36], or even a titanosaur ([6,75], LCDM here), but here identified as a probable MN (see main text for details). |
** Jobaria | Tiouaren Formation (Middle or Late Jurassic), Niger | Originally dated as Early Cretaceous by Sereno et al. [153], but more recently proposed as Middle Jurassic in age [145] |
Liubangosaurus | Napai Formation (Aptian), China | Identified as EU by Mo et al. [154], as a macronarian (LSDM here), a somphospondylan ([18]: [LSDM), and a saltasaurid ([18]: LSDM with implied weights; LCDM] and LCDM here). |
** Losillasaurus | Villar del Arzobispo Formation (late Tithonian-middle Berriasian), Spain | Identified as a diplodocoid by Casanovas et al. [155], but all subsequent analyses agree that this taxon is EU ([12,16,19,39,156], CSM here). |
*/** Oplosaurus | Wessex Formation (Barremian), UK | Identified as either EU or MN [1,36]. |
Sonorasaurus | Turney Ranch Formation (late Albian-early Cenomanian), USA | Recovered as MN by Royo-Torres [109], but other analyses place this taxon as a titanosauriform ([17,18], LCDM here). |
Tangvayosaurus | Grés Supérieurs Formation (Aptian-Albian), Laos | Recovered as MN by Royo-Torres [109], but all other analyses place this taxon in Somphospondyli ([17,18], LSDM and LCDM here). |
Tastavinsaurus | Forcall and Xert Formations (early Aptian), Spain | Recovered as MN by [14,15,19] and the CSM here, but most studies place this taxon within Titanosauriformes ([17,18,157], LSDM and LCDM here). |
** Turiasaurus | Villar del Arzobispo Formation (late Tithonian-middle Berriasian), Spain | Recovered as a titanosauriform by Mateus [158], but most studies have identified this taxon as EU ([19,109–111,159], CSM here). |
*Unnamed Dalton Wells taxon | Cedar Mountain Formation (Barremian), USA | Initially identified as a titanosaur (e.g. [160]), but more recently interpreted as MN ([161], PU pers. obs. 2013). |
*Unnamed Hell Canyon specimen | Lakota Formation (late Berriasian-Valanginian), USA | A Camarasaurus-like basal macronarian [162] |
Venenosaurus | Cedar Mountain Formation (Barremian), USA | Recovered as MN by Royo-Torres [109] and Carballido et al. [14,15], but most other analyses place this taxon as a brachiosaurid ([17–19], LCDM here) or somphospondylan ([122,157], CSM here). |
Wintonotitan | Winton Formation (Cenomanian-Turonian), Australia | Recovered as MN by Carballido et al. [14,15], but most other analyses place this taxon within Somphospondyli or Titanosauria (e.g. [1,18,19,80,163], CSM, LSDM and LCDM here). Poropat et al. [81] also presented additional character data that support inclusion of Wintonotitan within Somphospondyli. |
Xenoposeidon | Ashdown Formation (late Berriasian-Valanginian), UK | Phylogenetic analysis by Taylor and Naish [37] placed this taxon within Neosauropoda, and Upchurch et al. [36] and Mannion et al. [18] tentatively identified it as MN. The extreme incompleteness of the holotype (and only) specimen means that the affinities of this taxon remain uncertain. |
Xianshanosaurus | Mangchuan Formation (Cenomanian), China | Identified as a basal neosauropod by Lü et al. [136], but the phylogenetic analyses of Mannion et al. [18], and the LSDM/LCDM here, place this taxon within Somphospondyli or a more restricted clade within Titanosauria. |
‘*’ mark those taxa that are accepted here to be probable non-titanosauriform macronarians;
‘**’ mark those taxa that are accepted here to be probable non-neosauropod eusauropods.
Formation and age data were obtained from Fossilworks (http://fossilworks.org/), The Paleobiology Database (http://paleobiodb.org/#/) (both accessed on 15th August 2014), Upchurch et al. [36] and Mannion et al. [18]. Abbreviations: EU, non-neosauropod eusauropod; MN, non-titanosauriform macronarian.