Skip to main content
. 2015 Jun 3;10(6):e0125819. doi: 10.1371/journal.pone.0125819

Table 3. Summary of putative Early Cretaceous non-titanosauriform macronarians and other broad-toothed sauropods, according to previous studies.

Taxon Formation/Age/Country Identification
* Aragosaurus Villar del Arzobispo Formation (late Tithonian-middle Berriasian), Spain Identified as a titanosauriform by Canudo et al. [152] and D’Emic [17], but these opinions were not based on inclusion of Aragosaurus into a phylogenetic analysis. Recovered as MN in all analyses to date (e.g. [18,80,109,111]; LSDM and LCDM here).
Cedarosaurus Cedar Mountain Formation (Barremian-early Albian), USA Recovered as MN by Royo-Torres [109, but most other analyses place this taxon as a brachiosaurid [1,1719], LCDM here) or somphospondylan ([122], CSM here).
Chubutisaurus Cerro Barcino Formation (Aptian-Cenomanian), Argentina Recovered as a MN by Carballido et al. [15], but most other analyses place this taxon as a somphospondylan [14,1719], CSM here) or at least a titanosauriform (LCDM here).
Dongbeititan Yixian Formation (Barremian), China Recovered as MN in the LCDM analysis of Mannion et al. [18], but this is not supported by their LSDM analysis or the LCDM result here.
Euhelopus Mengyin Formation (Barremian-Aptian), China Recovered as MN by Carballido et al. [15], Carballido and Sander [19] and CSM Here, but most other analyses place this taxon within Somphospondyli ([10,13,17,18], LSDM and LCDM here) or at least a titanosauriform [14]
* Galveosaurus Villar del Arzobispo Formation (late Tithonian-middle Berriasian), Spain Recovered as EU by Royo-Torres and Upchurch [110] in part, and Royo-Torres et al. [111], but most studies support an MN position ([15,18,19], CSM, LSDM and LCDM here).
* Haestasaurus Hastings Beds Group (late Berriasian-Valanginian), UK Regarded as a titanosauriform [17], somphospondylan [18,36], or even a titanosaur ([6,75], LCDM here), but here identified as a probable MN (see main text for details).
** Jobaria Tiouaren Formation (Middle or Late Jurassic), Niger Originally dated as Early Cretaceous by Sereno et al. [153], but more recently proposed as Middle Jurassic in age [145]
Liubangosaurus Napai Formation (Aptian), China Identified as EU by Mo et al. [154], as a macronarian (LSDM here), a somphospondylan ([18]: [LSDM), and a saltasaurid ([18]: LSDM with implied weights; LCDM] and LCDM here).
** Losillasaurus Villar del Arzobispo Formation (late Tithonian-middle Berriasian), Spain Identified as a diplodocoid by Casanovas et al. [155], but all subsequent analyses agree that this taxon is EU ([12,16,19,39,156], CSM here).
*/** Oplosaurus Wessex Formation (Barremian), UK Identified as either EU or MN [1,36].
Sonorasaurus Turney Ranch Formation (late Albian-early Cenomanian), USA Recovered as MN by Royo-Torres [109], but other analyses place this taxon as a titanosauriform ([17,18], LCDM here).
Tangvayosaurus Grés Supérieurs Formation (Aptian-Albian), Laos Recovered as MN by Royo-Torres [109], but all other analyses place this taxon in Somphospondyli ([17,18], LSDM and LCDM here).
Tastavinsaurus Forcall and Xert Formations (early Aptian), Spain Recovered as MN by [14,15,19] and the CSM here, but most studies place this taxon within Titanosauriformes ([17,18,157], LSDM and LCDM here).
** Turiasaurus Villar del Arzobispo Formation (late Tithonian-middle Berriasian), Spain Recovered as a titanosauriform by Mateus [158], but most studies have identified this taxon as EU ([19,109111,159], CSM here).
*Unnamed Dalton Wells taxon Cedar Mountain Formation (Barremian), USA Initially identified as a titanosaur (e.g. [160]), but more recently interpreted as MN ([161], PU pers. obs. 2013).
*Unnamed Hell Canyon specimen Lakota Formation (late Berriasian-Valanginian), USA A Camarasaurus-like basal macronarian [162]
Venenosaurus Cedar Mountain Formation (Barremian), USA Recovered as MN by Royo-Torres [109] and Carballido et al. [14,15], but most other analyses place this taxon as a brachiosaurid ([1719], LCDM here) or somphospondylan ([122,157], CSM here).
Wintonotitan Winton Formation (Cenomanian-Turonian), Australia Recovered as MN by Carballido et al. [14,15], but most other analyses place this taxon within Somphospondyli or Titanosauria (e.g. [1,18,19,80,163], CSM, LSDM and LCDM here). Poropat et al. [81] also presented additional character data that support inclusion of Wintonotitan within Somphospondyli.
Xenoposeidon Ashdown Formation (late Berriasian-Valanginian), UK Phylogenetic analysis by Taylor and Naish [37] placed this taxon within Neosauropoda, and Upchurch et al. [36] and Mannion et al. [18] tentatively identified it as MN. The extreme incompleteness of the holotype (and only) specimen means that the affinities of this taxon remain uncertain.
Xianshanosaurus Mangchuan Formation (Cenomanian), China Identified as a basal neosauropod by Lü et al. [136], but the phylogenetic analyses of Mannion et al. [18], and the LSDM/LCDM here, place this taxon within Somphospondyli or a more restricted clade within Titanosauria.

‘*’ mark those taxa that are accepted here to be probable non-titanosauriform macronarians;

‘**’ mark those taxa that are accepted here to be probable non-neosauropod eusauropods.

Formation and age data were obtained from Fossilworks (http://fossilworks.org/), The Paleobiology Database (http://paleobiodb.org/#/) (both accessed on 15th August 2014), Upchurch et al. [36] and Mannion et al. [18]. Abbreviations: EU, non-neosauropod eusauropod; MN, non-titanosauriform macronarian.