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(A–C) Tomographic slices through T. thermophilus cells show invaginations in the outer membrane and large protein complexes crossing the periplasm (white arrowheads), which are associated with pili. Scale bar = 500 nm in A, 100 nm in B and C. (D) Volume rendering shows the distribution of pili (multi-coloured), protruding from the outer membrane (pale yellow). A concentric invagination of the outer membrane is indicated (white arrowheads).
