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Bidendate interactions of the arginine guanidinium group exemplified by (i) the carboxyl group of aspartate (trypsin–peptide complex, PDB code 1OX1) (left), (ii) the phosphoryl group of phosphotyrosine (SH2–pTyr, PDB code 4F5B) (middle),227 and (iii) atoms O6 and N7 of guanine (p53–DNA, PDB code 3TS8) (right).228
