Table 2.

Phosphorylated residues of keratins and kinases involved

Keratins	Phosphorylated residues	Kinases involved	References
K8	Ser-8	PKA, PKCɛ	(Akita et al., 2007; Ando et al., 1996)
	Ser-12	PKA	(Ando et al., 1996)
	Ser-23	PKA, PKCɛ	(Akita et al., 2007; Ando et al., 1996)
	Ser-33	PKA	(Ando et al., 1996)
	Ser-36	PKA	(Ando et al., 1996)
	Ser-42	PKA	(Ando et al., 1996)
	Ser-50	PKA	(Ando et al., 1996)
	Ser-73	JNK, PKCδ, MK2*	(He et al., 2002; Menon et al., 2010; Ridge et al., 2005)
	Ser-416	PKA	(Ando et al., 1996)
	Ser-423	PKA	(Ando et al., 1996)
	Ser-425	PKA	(Ando et al., 1996)
	Ser-431	ERK, JNK	(Busch et al., 2012; Park et al., 2011; Park et al., 2012)
	Not determined	AKT, AMPK, CAMK II, CK-Iα,	(Kuga et al., 2013; Loschke et al., 2015; Velasco et al., 1998; Yano et al., 1991)
K17	Ser-44	RSK1	(Pan et al., 2011)
	Not determined	US3	(Murata et al., 2002)
K18	Ser-33	PKCζ	(Sivaramakrishnan et al., 2009)
	Ser-52	MK2	(Menon et al., 2010)
	Not determined	AMPK	(Velasco et al., 1998)
K19	Ser-35	Not determined	(Zhou et al., 1999)
	Tyr-391	Src kinase	(Zhou et al., 2010)
K20	Ser-13	MK2, PKC*	(Menon et al., 2010; Zhou et al., 2006)

^*No evidence for phosphorylation of residue by indicated kinase but dependent on that.