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TreT-type trehalose synthase. (a) Natural reaction in trehalose synthesis direction starting from UDP-glucose and d-glucose; (b) active site residues (indicated in dark purple) surrounding the d-glucose moiety (light blue) in the acceptor binding pocket of P. horikoshii TreT [49]. Replacing the histidine at position 92 into an alanine via site-specific mutagenesis resulted in a 10% improvement of the activity on mannose as alternative acceptor monosaccharide.
