Niche partitioning within and among feeding guilds. NMDS of occurrence-based Bray–Curtis dissimilarity of samples from (A) all seven LMH species (adonis pseudo F6,285 = 59.0, R2 = 0.55, P ≤ 0.0001), (B) only grazers (pseudo F3,167 = 36.6, R2 = 0.40, P ≤ 0.0001), and (C) only nongrazers (pseudo F2,118 = 31.2, R2 = 0.35, P ≤ 0.0001). Symbols distinguish “grazers” (circles), “mixed feeders” (triangles), and “browsers” (squares).