Fig 1. Sample photoquadrats.

Sample photoquadrats acquired and annotated as part of long-term monitoring projects on shallow coral reefs (≤ 17-m depth). (A) Moorea, French Polynesia, acquired with strobes and framer (50 x 50 cm); (B) Palmyra, northern Line Islands, acquired using a manual white balance and framer (65 x 90 cm); (C) Nanwan Bay, Taiwan acquired with framer (35 x 35 cm) but neither strobes nor white balance; (D) Heron Reef, GBR, acquired without framer, strobes or white balance.