TABLE 1.

Ethogam Used for Behavior Quantification Separated into Behavior Categories Used for Statistical Analysis

Behavior Category	Behaviors	Definition
Hair-Directed	Hair chew	Mastication of either a loose or intact single strand or clump of hair from the individual’s own coat or that of another animal
	Hair-directed	Manual manipulation of an individual strand or clump of hair that may also involve placing the hair in the mouth and manipulating it with the lips without chewing
	Hair pull self	Forcefully removing hair from the individual’s own body in tufts or strands with hands or teeth
	Hair pull other	Forcefully removing hair from another individual in tufts or strands with hands or teeth
Biscuit-Directed	Eat biscuit	Biting and/or chewing biscuits
	Biscuit sort	Manual manipulation of biscuits, biscuit pieces, or crumbs including removing them from the food trough or picking through them on the ground
Forage-Directed	Forage-directed	Manual manipulation of forage with related chewing and consumption (note: the act of gathering and consuming forage occurred quickly and did not allow for the separation of forage manipulation from its chewing and consumption)
Active	Active	Any normal locomotion requiring more than two steps to be taken, as well as any changes in total body posture including sitting to standing or standing to sitting
Rest	Rest	Inactivity lasting for longer than 2 s
Object-Directed	Enrichment-directed	Oral or manual manipulation of any enrichment device in the cage
	Cage-directed	Any interaction with the cage such the manipulation of cage doors or parts
Social Behavior	Affiliative	Any gesture or facial expression directed toward the observer or another animal indicating sociability
	Aggressive	Any gesture or facial expression directed toward the observer or another animal intended to threaten or intimidate
	Submissive	Any gesture or facial expression directed toward the observer or another animal indicating deference
Groom Self	Groom self	Oral or manual manipulation of the individual’s own hair coat or body part without the plucking, removal, or chewing of the hair
Groom Other	Groom other	Oral or manual manipulation of another animal’s hair coat or body part without the plucking, removal, or chewing of the hair
Tension Related	Scratch	Brusque, stroking movement of the hand in which the fingers contact the skin or coat
	Yawn	Opening the mouth wide in a nonthreatening manner with a prolonged, deep inhalation and subsequent exhalation
Other	Other	Miscellaneous behaviors occurring too infrequently to allow for individual analysis