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. 2015 Jul 24;10(7):e0133720. doi: 10.1371/journal.pone.0133720

Table 1. Bioclimatic variables derived from temperature and precipitation.

Bio1 Annual mean temperature
Bio2 Mean diurnal range (mean monthly min. temp.—mean monthly max. temp.)
Bio3 Isothermality (Bio2/Bio7)
Bio4 Temperature seasonality (SD)
Bio5 Maximum temperature of the warmest month
Bio6 Minimum temperature of the coldest month
Bio7 Annual temperature range (Bio5-Bio6)
Bio8 Mean temperature of the wettest quarter
Bio9 Mean temperature of the driest quarter
Bio10 Mean temperature of the warmest quarter
Bio11 Mean temperature of the coldest quarter
Bio12 Annual precipitation
Bio13 Precipitation of the wettest month
Bio14 Precipitation of the driest month
Bio15 Precipitation seasonality (coefficient of variation)
Bio16 Precipitation of the wettest quarter
Bio17 Precipitation of the driest quarter
Bio18 Precipitation of warmest quarter
Bio19 Precipitation of coldest quarter