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a, The crystal structure of domain 4 (purple ribbons) from PA prepore (PDB ID: 1TZO) fits the cryoEM map (grey surface) of PA pore with a good agreement (cross-correlation coefficient: 0.91). b, Translocation of domain 4 during the prepore to pore conversion. Domain 4 is shifted inwards as a rigid-body for 4 Å from the prepore conformation (dark cyan) to the pore conformation (purple).
