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A proposed model for H2B monoubiquitination mediated by OsHUB1 and OsHUB2 during anther development in rice. In this model, OsHUB1 and OsHUB2 form a heterotetramer and recruit E2s to UDT1 and OsCP1 chromatin, leading to transfer of an ubiquitin molecule to H2B possibly at Lys-143. H2Bub1 formation enhances H3K4me2, together promoting transcription initiation. The fine-tuning of gene expression of UDT1/OsCP1 is important for late anther development of rice.
