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Early loss of centromere cohesion in pans1 during meiosis II. Acid spreads of male meiotic chromosomes stained with DAPI. a-d wild type, e-i pans1. a,e Normal metaphase I. b, f Normal anaphase I. c Normal metaphase II d Normal anaphase II. g Early loss of cohesion in pans1 meiosis II prior to metaphase II. h Random movement of chromosomes in meiosis II in pans1. i pans1 defective metaphase I showing bipolar attachment (arrowheads) and univalents (*) in a subset of chromosomes
