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Overall architecture of Complex I. The membrane domain is from E. coli Complex I, PDB file 3rko, and the peripheral domain is from T. thermophilus Complex I, PDB code 4hea. FMN in the peripheral arm is in red, and nine iron sulfur clusters are in orange. Two broken helices in each subunit LMN are indicated as TM7 in red and TM 12 in brown.
