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Proportion of bees extending the proboscis to a previously neutral odour following pairing with different unconditioned stimuli. For trials 1–3, the US was either 30% pollen (w/w) solution (Black triangles, forward paired, n = 34, White diamonds, backward paired, n = 34) or 30% sucrose (w/w) solution (Black squares, n = 32). All bees received 30% sucrose on trials 4–6. The US was applied to both the antennae and proboscis.
