Table 2.

Regional (γ-diversity) species and trait (combined and single) diversity

	γ-Diversity	% α	% β
Species diversity	5.80	50	50
Multi-trait diversity	1.87	84	16
Body length	1.26	94	6
Antenna/body length ratio	1.26	96	4
Life form	1.55	85	15
Moisture preference	1.44	94	6
Habitat width	1.35	94	6

% α and β give the proportional contributions of the local (α-diversity) and between-site turnover (β-diversity) components to the γ-diversity

Species diversity was calculated using the Simpson diversity index, trait diversity using Rao Q, and multi-trait diversity using all five traits together (see text for details). β-Diversity was calculated by additive partitioning