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Standard deviation in end boldness score. The standard deviation of boldness index within a colony at the end of the experiment significantly varied among treatments when the keystone individual (a) was removed after spending a variable amount of time within the colony (5, 10 or 20 days) prior to removal (x-axis), but not when a paired control shy individual (b) was removed after spending a variable amount of time within the colony (5, 10 or 20 days).
