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. 2015 Sep 15;112(37):11415–11422. doi: 10.1073/pnas.1501070112

Table 1.

Some theories of species diversity

Theory Author, date Ref(s).
Nonmechanistic
 Broken stick MacArthur, 1957 1
 Log normal Preston, 1962 2
 Log series Whittaker, 1965 3
 Neutral theory Hubbell, 1979, 2001 4, 5
Heterogeneity
 Intermediate disturbance Connell, 1978 6
 Lottery competition Chesson and Warner, 1981 7
 Spatial heterogeneity Tilman and Pacala, 1993 8
 Dispersal limitation Tilman, 1994 9
 Winner by forfeit Hurtt and Pacala, 1995 10
 Individual variation Clark, 2010 11
Bottom-up
 Resource competition Lotka, 1925 12
 Resource competition Volterra, 1926 13
 Limiting similarity Hutchinson, 1959 14
 Habitat complexity MacArthur and MacArthur, 1961 15
 Immigration vs. extinction MacArthur and Wilson, 1967 16
 Resource gradients Tilman, 1982 17
 Productivity Brown, 2014 18
Top-down
 Keystone predation Paine, 1966 19
 Escape in space Janzen, 1970 20
 Escape in space Connell, 1971 21
 Predation/competition Menge and Sutherland, 1976 22
 Pathogens Givnish, 1999 23
 Predation/competition Chesson and Kuang, 2008 24