Table 2.
Morphometry. Measurements in mm. Shell measures: AH; AW; BWW; SH; SW; W; statistics: CV; max; min; SD. First line of new species contains measurements of holotypes. Note that the holotype was only in case of Leiorhagium clandestinum included in the descriptive statistics. Numbers of whorls were only counted in the new species as this parameter was not used in the statistical analyses.
| New species | |||||||
| SH | SW | AH | AW | BWW | SH/SW | W | |
| Hemistomia andreae sp. n. (N=20) | 2.70 | 1.25 | 0.90 | 0.87 | 1.08 | 2.17 | 5.4 |
| min | 2.40 | 1.10 | 0.80 | 0.75 | 0.97 | 2.00 | 4.50 |
| max | 2.78 | 1.28 | 0.93 | 0.91 | 1.08 | 2.35 | 5.50 |
| mean | 2.60 | 1.18 | 0.85 | 0.82 | 1.02 | 2.20 | 5.14 |
| median | 2.60 | 1.17 | 0.85 | 0.82 | 1.01 | 2.23 | 5.25 |
| SD | 0.11 | 0.05 | 0.04 | 0.04 | 0.03 | 0.11 | 0.28 |
| CV | 4.40 | 3.93 | 4.23 | 4.54 | 2.77 | 4.94 | 5.49 |
| Leiorhagium adioincola sp. n. NeCa 49 (N=20) | 2.29 | 1.24 | 0.88 | 0.87 | 1.09 | 1.84 | 4.50 |
| min | 2.10 | 1.16 | 0.83 | 0.83 | 1.04 | 1.71 | 4.13 |
| max | 2.42 | 1.31 | 0.96 | 0.96 | 1.15 | 1.90 | 4.75 |
| mean | 2.25 | 1.25 | 0.88 | 0.89 | 1.10 | 1.80 | 4.36 |
| median | 2.24 | 1.24 | 0.88 | 0.89 | 1.10 | 1.80 | 4.25 |
| SD | 0.09 | 0.04 | 0.04 | 0.03 | 0.03 | 0.05 | 0.18 |
| CV | 4.21 | 2.96 | 4.15 | 3.92 | 2.94 | 2.91 | 4.24 |
| Leiorhagium aremuum sp. n. (N=20) | 2.19 | 1.35 | 0.97 | 0.91 | 1.16 | 1.62 | 4.25 |
| min | 2.03 | 1.29 | 0.87 | 0.86 | 1.10 | 1.53 | 3.75 |
| max | 2.43 | 1.46 | 1.03 | 1.00 | 1.25 | 1.69 | 4.25 |
| mean | 2.19 | 1.35 | 0.94 | 0.92 | 1.16 | 1.62 | 4.03 |
| median | 2.15 | 1.35 | 0.93 | 0.92 | 1.17 | 1.62 | 4.00 |
| SD | 0.11 | 0.05 | 0.04 | 0.04 | 0.04 | 0.04 | 0.15 |
| CV | 4.92 | 4.06 | 4.76 | 4.54 | 3.77 | 2.71 | 3.78 |
| Leiorhagium clandestinum sp. n. (N=4) | 2.49 | 1.32 | 0.94 | 0.95 | 1.16 | 1.91 | 4.50 |
| min | 2.23 | 1.26 | 0.89 | 0.88 | 1.07 | 1.77 | 4.25 |
| max | 2.49 | 1.32 | 0.94 | 0.95 | 1.16 | 1.91 | 4.50 |
| mean | 2.38 | 1.28 | 0.91 | 0.92 | 1.10 | 1.86 | 4.41 |
| median | 2.41 | 1.27 | 0.90 | 0.93 | 1.09 | 1.89 | 4.44 |
| SD | 0.11 | 0.03 | 0.02 | 0.03 | 0.04 | 0.06 | 0.12 |
| CV | 4.83 | 2.30 | 2.68 | 3.44 | 4.00 | 3.52 | 2.89 |
| Leiorhagium neteae n. sp. (N=18) | 2.07 | 1.12 | 0.75 | 0.77 | 0.91 | 1.84 | 4.50 |
| min | 1.85 | 0.97 | 0.65 | 0.70 | 0.82 | 1.76 | 4.25 |
| max | 2.23 | 1.17 | 0.79 | 0.80 | 0.95 | 2.01 | 5.00 |
| mean | 2.05 | 1.09 | 0.73 | 0.75 | 0.88 | 1.88 | 4.46 |
| median | 2.04 | 1.10 | 0.73 | 0.75 | 0.87 | 1.88 | 4.38 |
| SD | 0.12 | 0.05 | 0.03 | 0.03 | 0.03 | 0.07 | 0.19 |
| CV | 6.05 | 4.82 | 4.71 | 4.69 | 3.99 | 3.72 | 4.25 |
| Material for comparisons | |||||||
| SH | SW | AH | AW | BWW | SH/SW | ||
| Hemistomia cockerelli Types (N=20) | |||||||
| min | 2.58 | 1.18 | 0.88 | 0.83 | 1.03 | 2.05 | |
| max | 3.21 | 1.39 | 1.03 | 0.97 | 1.16 | 2.40 | |
| mean | 2.79 | 1.27 | 0.94 | 0.91 | 1.09 | 2.19 | |
| median | 2.74 | 1.25 | 0.93 | 0.90 | 1.09 | 2.18 | |
| SD | 0.17 | 0.06 | 0.04 | 0.04 | 0.05 | 0.09 | |
| CV | 6.20 | 4.91 | 4.31 | 4.52 | 4.23 | 4.36 | |
| Hemistomia cockerelli NeCa11 (N=20) | |||||||
| min | 2.20 | 1.06 | 0.77 | 0.73 | 0.94 | 1.93 | |
| max | 2.48 | 1.25 | 0.87 | 0.91 | 1.04 | 2.28 | |
| mean | 2.33 | 1.13 | 0.81 | 0.81 | 0.97 | 2.06 | |
| median | 2.32 | 1.12 | 0.80 | 0.80 | 0.96 | 2.03 | |
| SD | 0.08 | 0.04 | 0.03 | 0.04 | 0.02 | 0.10 | |
| CV | 3.49 | 3.70 | 3.36 | 4.60 | 2.36 | 4.75 | |
| Hemistomia cockerelli NeCa17 (N=20) | |||||||
| min | 2.35 | 1.16 | 0.83 | 0.83 | 1.04 | 1.96 | |
| max | 2.62 | 1.28 | 0.92 | 0.93 | 1.14 | 2.19 | |
| mean | 2.50 | 1.21 | 0.87 | 0.87 | 1.08 | 2.07 | |
| median | 2.51 | 1.21 | 0.87 | 0.88 | 1.08 | 2.07 | |
| SD | 0.07 | 0.04 | 0.03 | 0.03 | 0.02 | 0.07 | |
| CV | 2.90 | 3.20 | 3.16 | 3.25 | 2.32 | 3.43 | |
| Hemistomia cockerelli NeCa21A (N=8) | |||||||
| min | 2.26 | 1.09 | 0.74 | 0.77 | 0.96 | 2.03 | |
| max | 2.74 | 1.23 | 0.89 | 0.87 | 1.08 | 2.38 | |
| mean | 2.49 | 1.17 | 0.84 | 0.83 | 1.03 | 2.12 | |
| median | 2.49 | 1.17 | 0.85 | 0.83 | 1.05 | 2.09 | |
| SD | 0.14 | 0.05 | 0.04 | 0.03 | 0.04 | 0.11 | |
| CV | 5.87 | 4.33 | 5.41 | 3.89 | 3.66 | 5.50 | |
| Hemistomia cockerelli NeCa36 (N=13) | |||||||
| min | 2.32 | 1.14 | 0.79 | 0.82 | 1.03 | 1.97 | |
| max | 2.64 | 1.23 | 0.91 | 0.91 | 1.12 | 2.14 | |
| mean | 2.43 | 1.18 | 0.85 | 0.85 | 1.06 | 2.05 | |
| median | 2.42 | 1.19 | 0.86 | 0.85 | 1.06 | 2.04 | |
| SD | 0.10 | 0.03 | 0.03 | 0.03 | 0.03 | 0.05 | |
| CV | 4.25 | 2.42 | 3.79 | 3.22 | 2.64 | 2.64 | |
| Hemistomia cockerelli NeCa54 (N=20) | |||||||
| min | 2.28 | 1.16 | 0.78 | 0.82 | 1.04 | 1.86 | |
| max | 2.63 | 1.31 | 0.96 | 0.93 | 1.14 | 2.14 | |
| mean | 2.47 | 1.23 | 0.87 | 0.88 | 1.09 | 2.00 | |
| median | 2.47 | 1.23 | 0.86 | 0.87 | 1.10 | 2.02 | |
| SD | 0.10 | 0.04 | 0.04 | 0.03 | 0.03 | 0.07 | |
| CV | 4.18 | 3.08 | 4.18 | 3.28 | 2.62 | 3.62 | |
| Hemistomia nyo NeCa35 (N=7) | |||||||
| min | 2.43 | 1.25 | 0.88 | 0.89 | 1.09 | 1.93 | |
| max | 2.75 | 1.34 | 0.96 | 0.96 | 1.15 | 2.08 | |
| mean | 2.62 | 1.30 | 0.92 | 0.92 | 1.12 | 2.01 | |
| median | 2.69 | 1.30 | 0.91 | 0.91 | 1.11 | 2.03 | |
| SD | 0.12 | 0.04 | 0.03 | 0.03 | 0.02 | 0.06 | |
| CV | 4.80 | 2.84 | 3.30 | 3.10 | 2.00 | 3.00 | |
| Leiorhagium ajie Types (N=6) | |||||||
| min | 2.35 | 1.31 | 0.93 | 0.94 | 1.12 | 1.61 | |
| max | 2.74 | 1.62 | 1.10 | 1.06 | 1.34 | 1.80 | |
| mean | 2.50 | 1.46 | 1.01 | 1.00 | 1.25 | 1.72 | |
| median | 2.43 | 1.46 | 1.01 | 1.00 | 1.27 | 1.70 | |
| SD | 0.16 | 0.12 | 0.07 | 0.05 | 0.08 | 0.07 | |
| CV | 6.50 | 8.31 | 6.95 | 4.88 | 6.32 | 4.12 | |
| Leiorhagium douii Types (N=20) | |||||||
| min | 1.87 | 0.98 | 0.68 | 0.68 | 0.86 | 1.84 | |
| max | 2.50 | 1.16 | 0.84 | 0.79 | 0.97 | 2.16 | |
| mean | 2.06 | 1.05 | 0.73 | 0.71 | 0.91 | 1.96 | |
| median | 2.02 | 1.06 | 0.72 | 0.71 | 0.91 | 1.95 | |
| SD | 0.14 | 0.04 | 0.04 | 0.02 | 0.03 | 0.08 | |
| CV | 7.04 | 4.02 | 5.11 | 3.51 | 3.21 | 4.23 | |
| Leiorhagium kavuneva Types (N=20) | |||||||
| min | 2.17 | 1.17 | 0.78 | 0.82 | 1.02 | 1.77 | |
| max | 2.42 | 1.32 | 0.94 | 0.93 | 1.13 | 1.93 | |
| mean | 2.31 | 1.26 | 0.88 | 0.88 | 1.07 | 1.84 | |
| median | 2.33 | 1.25 | 0.89 | 0.88 | 1.07 | 1.85 | |
| SD | 0.07 | 0.04 | 0.04 | 0.03 | 0.03 | 0.05 | |
| CV | 3.24 | 3.16 | 4.52 | 3.15 | 2.87 | 2.58 | |
| Leiorhagium kavuneva NeCa15B (N=20) | |||||||
| min | 2.20 | 1.21 | 0.84 | 0.88 | 1.07 | 1.76 | |
| max | 2.46 | 1.31 | 0.94 | 0.98 | 1.20 | 1.97 | |
| mean | 2.34 | 1.27 | 0.90 | 0.92 | 1.12 | 1.84 | |
| median | 2.35 | 1.28 | 0.91 | 0.92 | 1.12 | 1.83 | |
| SD | 0.07 | 0.03 | 0.03 | 0.03 | 0.03 | 0.06 | |
| CV | 3.14 | 2.30 | 3.31 | 3.00 | 2.54 | 3.14 | |
| Leiorhagium kavuneva NeCa29 (N=20) | |||||||
| min | 2.17 | 1.20 | 0.85 | 0.85 | 1.06 | 1.76 | |
| max | 2.54 | 1.36 | 1.00 | 0.99 | 1.17 | 1.97 | |
| mean | 2.35 | 1.28 | 0.91 | 0.93 | 1.12 | 1.83 | |
| median | 2.34 | 1.27 | 0.90 | 0.93 | 1.13 | 1.82 | |
| SD | 0.10 | 0.04 | 0.04 | 0.04 | 0.03 | 0.05 | |
| CV | 4.27 | 3.12 | 4.50 | 3.93 | 2.91 | 2.62 | |
| Leiorhagium monachum Types (N=3) | |||||||
| min | 2.07 | 1.04 | 0.72 | 0.69 | 0.88 | 1.88 | |
| max | 2.18 | 1.10 | 0.82 | 0.78 | 0.97 | 2.00 | |
| mean | 2.11 | 1.08 | 0.76 | 0.74 | 0.92 | 1.96 | |
| median | 2.07 | 1.09 | 0.76 | 0.75 | 0.92 | 1.99 | |
| SD | 0.07 | 0.03 | 0.05 | 0.05 | 0.05 | 0.07 | |
| CV | 3.36 | 3.19 | 7.25 | 6.89 | 5.51 | 3.67 | |
| Leiorhagium montfaouense Types (N=10) | |||||||
| min | 1.80 | 1.03 | 0.68 | 0.64 | 0.83 | 1.76 | |
| max | 2.30 | 1.16 | 0.81 | 0.77 | 0.99 | 1.99 | |
| mean | 2.01 | 1.08 | 0.73 | 0.70 | 0.90 | 1.87 | |
| median | 2.02 | 1.05 | 0.72 | 0.68 | 0.89 | 1.86 | |
| SD | 0.15 | 0.05 | 0.05 | 0.04 | 0.06 | 0.09 | |
| CV | 7.73 | 4.89 | 6.40 | 6.29 | 6.87 | 4.79 | |