Table 2.
URI table for morphometric characters and Hymenoptera-specific terminology of morphological statements used in descriptions, identification key, and diagnoses are mapped to classes in phenotype-relevant ontologies.
| Abbr. | Label | Class genus differentia definition | Comments | uri |
|---|---|---|---|---|
| CL | maximum cephalic length in median view | The median anatomical line that extends between the posterior margin of the cranium and the distal margin of the clypeus in frontal view. | The maximum cephalic length in median view is not equivalent to the maximum cephalic size that extends between the posterior cranial margin and the distal clypeal line. The head must be carefully tilted to the position with the true maximum. Excavations of hind vertex and/or clypeus reduce CL (Fig. 1A). | http://purl.obolibrary.org/obo/HAO_0002331 |
| CW | head width | The anatomical line that is the longest horizontal diameter of the cranium in frontal view. | The head width is the largest distance between the lateral margins of the compound eyes measured in frontal view (Fig. 1A). | http://purl.obolibrary.org/obo/HAO_0002268 |
| CWb | dorsal head width | The anatomical line between the intersections of the cranium contour line and dorsal head line in frontal view. | The dorsal head width is the maximum width of head capsule without the compound eyes that is measured just posterior of the eyes in frontal view (Fig. 1A). | http://purl.obolibrary.org/obo/HAO_0002314 |
| Cdep | median clypeal notch depth | The anatomical line that is between the distal clypeal line and the proximalmost point of the distal clypeal notch in frontal view. | http://purl.obolibrary.org/obo/HAO_0002333 | |
| EL | maximum diameter of compound eye | The longest diameter of the eye. | http://purl.obolibrary.org/obo/HAO_0002326 | |
| FRS | frontal carina line | The transverse torular line that extends between the frontal carinae. | Distance of the frontal carinae immediately caudal of the posterior intersection points between frontal carinae and the torular lamellae. If these dorsal lamellae do not laterally surpass the frontal carinae, the deepest point of scape corner pits may be taken as reference line. These pits take up the inner corner of scape base when the scape is fully switched caudad and produce a dark triangular shadow in the lateral frontal lobes immediately posterior of the dorsal lamellae of scape joint capsule (Fig. 1B). | http://purl.obolibrary.org/obo/HAO_0002323 |
| ML | Weber length | The anatomical line that connects the global minima of the contour line of the pronotal slope in lateral view when the specimen is rotated until the contour line becames as symmetric as possible and the posteriormost point of the propodeal lobe. | Preferentially measured in lateral view; if the transition point is not well defined, use dorsal view and take the centre of the dark-shaded borderline between pronotal slope and pronotal shield as anterior reference point. In gynes: length from distalmost point of propodeal lobe to the most distant point of steep anterior pronotal face (Fig. 1E). | http://purl.obolibrary.org/obo/HAO_0002309 |
| MPST | maximum spiracle distance of propodeum | The anatomical line that connects the center of the propodeal spiracle with the posteriormost point of the propodeal lobe in lateral view. | Maximum distance from the center of the propodeal stigma to the anterioventral corner of the ventrolateral margin of the metapleuron (Fig. 1F). | http://purl.obolibrary.org/obo/HAO_0002334 |
| MW | mesosoma width | The longest width of the pronotum in dorsal view. | Mesosoma width. In workers MW is defined as the longest width of the pronotum in dorsal view excluding the pronotal spines (Fig. 1D). | http://purl.obolibrary.org/obo/HAO_0002335 |
| NOL | length of petiolar node | The anatomical line that is the shortest between the center of the petiolar spiracle and the posterior margin of the petiole in lateral view. | Length of the petiolar node. Measured in lateral view from the centre of petiolar spiracle to posterodorsal corner of caudal cylinder. Do not erroneously take as reference point the dorso-caudal corner of the helcium, which is sometimes visible (Fig. 1F). | http://purl.obolibrary.org/obo/HAO_0002336 |
| NOH | maximum height of petiolar node | The anatomical line that is the longest between the dorsal margin of the petiole and the posterior petiolar distance and perpendicular to the posterior petiolar distance. | http://purl.obolibrary.org/obo/HAO_0002327 | |
| NSTI | apical petiolar spine distance | The anatomical line between the distal ends of the anterodorsal spines of the petiolar node. | If spine tips are rounded or thick take the centers of spine tips as reference points (Fig. 1C). | http://purl.obolibrary.org/obo/HAO_0002338 |
| PEH | maximum petiole height | The anatomical line that is the longest between the ventral margin of the petiole and the dorsal margin of the petiole and is perpendicular to the ventral margin of the petiole in lateral view. | http://purl.obolibrary.org/obo/HAO_0002328 | |
| PEL | diagonal petiolar length | The anatomical line that extends between the distalmost point of the subpetiolar process and the global minima of the contour line of the dorsal region of the posterior petiolar constriction in lateral view when the specimen is rotated until the contour line became as symmetric as possible. | Fig. 1E. | http://purl.obolibrary.org/obo/HAO_0002317 |
| PEW | petiole width | The maximum width of the petiole in dorsal view. | Anterodorsal spines of the petiolar node are not considered (Fig. 1D). | http://purl.obolibrary.org/obo/HAO_0002339 |
| PoOC | postocular distance | The median anatomical line of the cranium that is the longest between the dorsal margin of the cranium and the dorsal head width. | Use a cross-scaled ocular micrometer and adjust the head to the measuring position of CL. Caudal measuring point: median occipital margin; frontal measuring point: median head at the level of the posterior eye margin (Fig. 1A). | http://purl.obolibrary.org/obo/HAO_0002340 |
| PPL | postpetiole length | The longest anatomical line that is perpendicular to the posterior margin of the postpetiole in lateral view and is between the posterior postpetiolar margin and the anterior postpetiolar margin. | Fig. 1F | http://purl.obolibrary.org/obo/HAO_0002341 |
| PPW | postpetiole width | The maximum width of the postpetiole in dorsal view. | Fig. 1D | http://purl.obolibrary.org/obo/HAO_0002342 |
| PSTI | apical distance of pronotal spines | The anatomical line between the distal ends of the pronotal spines. | If spine tips are rounded or thick take the centers of spine tips as reference points (Fig. 1D). | http://purl.obolibrary.org/obo/HAO_0002345 |
| SL | scape length | The proximodistal anatomical line of the scapal area distal to the radicle. | Maximum straight line scape length excluding the radicle (Fig. 1A). | http://purl.obolibrary.org/obo/HAO_0002346 |
| SPBA | minimum spine distance | The shortest anatomical line between the lateral margins of the propodeal spines. | This should be measured in anterodorsal view, since the wider parts of the ventral propodeum do not interfere with the measurement in this position. If the lateral margins of spines diverge continuously from the tip to the base, a smallest distance at base is not defined. In this case, SPBA is measured at the level of the bottom of the interspinal meniscus (Fig. 1C). | http://purl.obolibrary.org/obo/HAO_0002347 |
| SPST | spine length | The anatomical line between the center of the propodeal spiracle and the distal end of the propodeal spine. | Spine length. Distance between the centre of propodeal stigma and spine tip. The stigma centre refers to the midpoint defined by the outer cuticular ring but not to the centre of real stigma opening that may be positioned eccentrically (Fig. 1F). | http://purl.obolibrary.org/obo/HAO_0002348 |
| SPTI | apical spine distance | The anatomical line between the distal ends of the propodeal spines. | If spine tips are rounded or truncated, the centres of spine tips are taken as reference points (Fig. 1C). | http://purl.obolibrary.org/obo/HAO_0002319 |
| anterior pronotal slope | The concave area anteriorly on the mesosoma that accommodates the posterior area of the cranium. | http://purl.obolibrary.org/obo/HAO_0002311 | ||
| anterior setal pit | The anteriormost setal pit on the dorsal side of the petiole. | http://purl.obolibrary.org/obo/HAO_0002312 | ||
| caudal cylinder | The petiolar area posterior to the posterior petiolar constriction. | http://purl.obolibrary.org/obo/HAO_0002318 | ||
| cranial scrobe of the pronotum | The scrobe on the pronotum that accommodates the posterior surface of the cranium. | http://purl.obolibrary.org/obo/HAO_0002343 | ||
| distal clypeal line | The anatomical line that is perpendicular to the median anatomical line and is the tangent at the distalmost point(s) of the clypeus in frontal view. | http://purl.obolibrary.org/obo/HAO_0002316 | ||
| dorsal head line | The anatomical line between the posteriormost (dorsalmost) points of compound eyes in frontal view. | http://purl.obolibrary.org/obo/HAO_0002315 | ||
| dorsal petiolar scrobe | The scrobe that is dorsal to the propodeal foramen and accommodates the proximodorsal area of the petiole. | http://purl.obolibrary.org/obo/HAO_0002313 | ||
| external area of the scape | The area of the scape that faces away from the cranial surface in fully caudal scape position. | http://purl.obolibrary.org/obo/HAO_0002320 | ||
| eye | The compound organ that is composed of ommatidia. | http://purl.obolibrary.org/obo/HAO_0000217 | ||
| facial area of the scape | The area of the scape that faces the cranium surface when the scape is in fully flexed position. | http://purl.obolibrary.org/obo/HAO_0002321 | ||
| frontal carina | The carina that extends along the lateral margin of the intertorular area (median margin of the antennal foramen) towards the vertex. | http://purl.obolibrary.org/obo/HAO_0001533 | ||
| frontal carina line | The transverse torular line that extends between the frontal carinae. | http://purl.obolibrary.org/obo/HAO_0002323 | ||
| lateral carina of clypeus | The carina that extends between the ventral (anterior) margin of the antennal foramen to the apical clypeal margin. | http://purl.obolibrary.org/obo/HAO_0002324 | ||
| margin | The line that delimits the periphery of an area. | http://purl.obolibrary.org/obo/HAO_0000510 | ||
| median clypeal notch | The median notch that is on the distal clypeal margin. | http://purl.obolibrary.org/obo/HAO_0002332 | ||
| mesosoma | The anatomical cluster that is composed of the prothorax, mesothorax and the metapectal-propodeal complex. | http://purl.obolibrary.org/obo/HAO_0000576 | ||
| Weber length | The anatomical line that connects the global minima of the contour line of the pronotal slope in lateral view when the specimen is rotated until the contour line becames as symmetric as possible and the posteriormost point of the propodeal lobe. | http://purl.obolibrary.org/obo/HAO_0002309 | ||
| petiolar scrobe | The scrobe that is located ventrally of the propodeal foramen and accommodates the proximal area of the petiole. | http://purl.obolibrary.org/obo/HAO_0002265 | ||
| pronotal spine | The spine that is located at the dorsolateral edge of the cranial scrobe of the pronotum. | http://purl.obolibrary.org/obo/HAO_0002344 | ||
| pronotum | The notum that is located in the prothorax. | http://purl.obolibrary.org/obo/HAO_0000853 | ||
| scape | The antennal segment that is proximal to the pedicel and is connected to the head via the radicle. | http://purl.obolibrary.org/obo/HAO_0000908 | ||
| scrobe | The area that is impressed and is for the reception or concealment of another sclerite. | http://purl.obolibrary.org/obo/HAO_0000912 | ||
| setal angle | The angle of the proximodistal axis of the seta to the contour line of the bodypart where the seta is located. | http://purl.obolibrary.org/obo/HAO_0002330 | ||
| setal line | The row that is composed of setae. | http://purl.obolibrary.org/obo/HAO_0000903 | ||
| setal pit | The impression with a centered sensillum trichodeum. | http://purl.obolibrary.org/obo/HAO_0001958 | ||
| spine | The process that lacks non-sclerotised ring at the base. | http://purl.obolibrary.org/obo/HAO_0000949 | ||
| spiracle | The anatomical cluster that is composed of the distal end of the trachea and the margin of the sclerite or conjunctiva surrounding the spiracular opening. | http://purl.obolibrary.org/obo/HAO_0000950 | ||
| transverse torular line | The anatomical line that is tangential to the posteriormost points of the antennal rims. | http://purl.obolibrary.org/obo/HAO_0002322 | ||
| width | A 1-D extent quality which is equal to the distance from one side of an object to another side which is opposite. | http://purl.obolibrary.org/obo/HAO_0002308 |