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Mating behavior of Bursaphelenchus okinawaensis and B. xylophilus. The images show the stages of (A) pre-mating behavior of B. okinawaensis; (B) spicule insertion behavior of B. okinawaensis; (C) premating behavior of B. xylophilus; and (D) spicule insertion behavior of B. xylophilus. M, male; H, hermaphrodite; F, female. White arrows indicate the position of male spicule inserted into vulva.
