Overexpression of neither UGT84A1 nor UGT84A3 rescues the HCE deficiency of the fah1 mutant. Shown are steady-state transcript levels of UGT84A1 and UGT84A3 (A), ferulate:UDP-Glc glucosyltransferase activity (B), and hydroxycinnamoylmalate content (C) of 3-week-old whole rosettes of wild-type plants, fah1 plants, and T1 progeny of fah1 plants transformed with either a UGT84A1 or UGT84A3 overexpression construct. Transcript abundance in A is shown relative to that of the control gene At1g13320. Graphs in A and C show means ± sd of three biological replicates of individual plants, whereas those in B show means ± sd of three technical replicates of pooled samples.